(please email me for reprints; publications in Google Scholar)

Taram, M., Borah, D., Krishna, N., Pradeep, A.K., Amrutha, A. & Hughes, M. (2020). Begonia oyuniae (Begonia sect. Monophyllon, Begoniaceae), a remarkable new species from Northeast India. Gardens' Bulletin Singapore 72(1): 109-115.

Hughes, M., Girmansyah, D., Randi, A. & Ningsih, H.N.R. (2020). Eleven new records, three new species and an updated checklist of Begonia from Kalimantan, Indonesia. Gardens' Bulletin Singapore 72(1): 33-58.

Phutthai, T., Hughes, M. & Sridith, K. (2020). Begoniaceae. Flora of Thailand 14(3): 359-431.

Lanorsavanh, S., Hughes, M., Souvannkhoummane, K. & Lamxay, V. (2020). Begonia phouchomvoyensis (Begoniaceae), a new species from Lao PDR. Taiwania 65(1): 37-40.

Atkins, H.J., Bramley, G.L., Johnson, M.A., Kartonegoro, A., Nishii, K., Kokubugata, G., Nishii, K., Kokunugata, G., Möller, M. & Hughes, M. (2019). A molecular phylogeny of Southeast Asian Cyrtandra (Gesneriaceae) supports an emerging paradigm for Malesian plant biogeography. Frontiers of Biogeography.

Forrest, L.L., Hart, M.L., Hughes, M., Wilson, H.P., Chung, K.-F., Tseng, Y.-H. & Kidner, C.A. (2019). The limits of Hyb-Seq for herbarium specimens: Impact of preservation techniques. Front. Ecol. Evol. 7: 439.

Marasinghe, L.D.K., Rubasinghe, S.C.K., & Hughes, M. (2019). Rediscovery of the Sri Lanka endemic Amomum nemorale from Kanneliya Forest Reserve, Southern Province. NeBIO 10(4).

Wilson, H.P., Paul, O. & Hughes, M. (2019). Begonia maguniana (Begoniaceae, Begonia sect. Oligandrae), a new species from New Guinea. Edinburgh Journal of Botany doi: 10.1017/S0960428619000283 Read only pdf

Ardi, W.H., Girmansyah, D., Lin, C.W. & Hughes, M. (2019). Two new species of Begonia (Begoniaceae) from Borneo. Phytotaxa 407 (1): 22-28.

Girmansyah, D., Susila & Hughes, M. (2019). A revision of Begonia section. Petermannia on Sumatra, Indonesia. Phytotaxa 407(1): 79-100.

Hughes, M., Aung, M.M. & Armstrong, K. (2019). An updated checklist and a new species of Begonia (B. rheophytica) from Myanmar. Edinburgh Journal of Botany pdf

Borg, D., Richardson, J.E., Harris, D.J., Gautier, L., Hughes, M. & Mackinder, B. (2019). Phylogeny of two African genera of Sapotaceae - Englerophytum and Synsepalum. Edinburgh Journal of Botany pdf

Naive, M.A.K., Alejandro, G.J.D. & Hughes, M. (2018). Taxonomic notes on the Philippine endemic Begonia colorata (Begoniaceae, section Petermannia). Phytotaxa 381 (1): 127–131.

Hughes, M., Peng, C.-I, Liu, Y., Thomas, D.C., Phutthai, T., Rubite, R.R, Girmansyah, D., Ardi, W.H., Tanaka, N., Mui, C.Y. & Tan, J. (2018). Species Portraits. In: Hughes, M. & Peng, C.-I (Eds.) Asian Begonia: 300 Species Portraits. KBCC Press, Taipei, Taiwan & RBGE, Edinburgh, Scotland.

Adri, W.H. & Hughes, M. (2018). Two new species of Begonia from Sumatra. Edinburgh Journal of Botany doi: 10.1017/S0960428618000136

Kiew, R., Julia, S., Ling, C.Y., Randi, A., Girmansyah, D., Hughes, M. (2018). Taxonomic status of Begonia promethea (sect. Petermannia, Begoniaceae) in Borneo. Gardens' Bulletin Singapore 70(1): 155-161.

Moonlight, P.W., Ardi, W.H., Padilla, L.A., Chung, K.-F., Fuller, D., Girmansyah, D., Hollands, R., Jara-Muñoz, A., Kiew, R., Leong, W.C., Liu, Y., Mahardika, A., Marasinghe, L.D.K., O’Connor, M., Peng, C.-I., Pérez, Á.J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R.R., Julia, S., Scherberich, D., Shui, Y.-M., Tebbitt, M.C., Thomas, D.C., Wilson, H.P., Zaini, N.H., Hughes, M., (2018). Dividing and conquering the fastest growing genus: Towards a natural sectional classification of the mega-diverse genus Begonia (Begoniaceae). Taxon 67, 267–323

Hughes, M., Peng, C.-I, Lin, C.W., Rubite, R.R., Blanc, P., Chung, K-F. (2018) Chloroplast and nuclear DNA exchanges among Begonia sect. Baryandra species (Begoniaceae) from Palawan Island, Philippines, and descriptions of five new species. PLOSone 0194877

Rubite, R.R., Peng, C.-I, Chung, K-F., Line, C.W., Evangelista, L.T., Tandang, D.N., Callado, J.R. & Hughes, M. (2018). Three new species of Begonia (section Baryandra, Begoniaceae) from Luzon Island, the Philippines. Phytotaxa 347(3): 201-212.

Lý, N.S., Peng, C.-I & Hughes, M. (2018). Begonia lecongkietii (sect. Petermannia), a new species from Mount Dầu, South Central Coast Region, Vietnam. Edinburgh Journal of Botany doi: 10.1017/S0960428618000033 pdf

Gagul, J., Sands, M.J.S., Gideon, O. & Hughes, M. (2018). A revision of Begonia sect. Symbegonia on New Guinea. Edinburgh Journal of Botany doi: 10.1017/S096042861800001X pdf

Camfield, R. & Hughes, M. (2018). A revision and one new species of Begonia (Begoniaceae, Cucurbitales) in northeast India. European Journal of Taxonomy 369: 1-116.

Peng, C.-I, Rubite, R., Lin, C.W., Hughes, M., Kono, Y. & Chung, K.F. (2017). Three new species of Begonia sect. Baryandra from Panay Island, Philippines. Botanical Studies 58: 28 DOI 10.1186/s40529-017-0182-x

Phutthai, T. & Hughes, M. (2017). Four new species of Begonia (Begoniaceae) from Thailand. Edinburgh Journal of Botany. //

Tebbitt, M.C., Hughes, M., Perez, A.J. & Moonlight, P.W.(2017). Taxonomy of the Begonia tiliifolia group, including descriptions of two new species. Edinburgh journal of Botany. //

Moonlight, P.W., Hughes, M. & Tebbitt, M.C. (2017). Taxonomy of Begonia albomaculata and description of two new species endemic to Peru. Edinburgh Journal of Botany. //

Phutthai, T. & Hughes, M. (2017). A new species of Begonia sect. Parvibegonia (Begoniaceae) from Thailand and Myanmar. Blumea 62: 26-28.

Souvannakhoummane, K., Hughes, M. & Lanorsavanh, S. (2016). Begonia lamxayiana Souvann. (Begoniaceae): a new species from Lao PDR. Thai Journal of Botany 8(1): 1-5.

Phutthai, T. & Hughes, M. (2016). A new species and a new record in Begonia sect. Platycentrumfrom Thailand. Gardens' Bulletin Singapore 68(1): 99-107.

Hughes, M., Girmansyah, D. & Handoyo, W.I. (2015). Further discoveries in the ever expanding genus Begonia (Begoniaceae): fifteen new species from Sumatra. European Journal of Taxonomy 167: 1-40.

Nishii, K., M. Hughes, M. Briggs, E. Haston, F. Christie, M.J. deVilliers, T. Hanekom, W.G. Roos, D.U. Bellstedt, M. Möller. (2015) Streptocarpus redefined to include all Afro-Malagasy Gesneriaceae: Molecular phylogenies prove congruent with geography and cytology and uncovers remarkable morphological homoplasies. Taxon 64(6): 1243-1274.

Pennington, R.T., Hughes, M. & Moonlight, P.W. (2015). The origins of tropical rainforest hyperdiversity. Trends in Plant Science

Wang J., Chen W.H., Hughes M., Yang Z.Z. & Shui Y.M. (2015). Additional notes on the Begonia sect. Petermannia (Begoniaceae) from China. Plant Diversity and Resources 37 (5): 563–568.

Rubite, R.R., Hughes, M., Blanc, P., Chung, K.F., Yang, H.A., Kono, Y., Alejandro, G.J.D., De Layola, L.B., Virata, A.G.N. and Peng, C.-I. (2015). Three new species of Begonia endemic to the Puerto Princesa Subterranean River National Park, Palawan. Botanical Studies 56: 19.

Hughes, M., Rubite, R., Blanc, P., Fang, K.-F. & Peng, C.-I. (2015) The Miocene to Pleistocene colonization of the Philippine archipelago by Begonia sect. Baryandra (Begoniaceae). American Journal of Botany (online early).

Hughes, M., Barber, S., Heatabun, C. & Gagul, J. (2015). Begonia yapenensis (sect. Symbegonia, Begoniaceae), a new species from Papua, Indonesia. European Journal of Taxonomy, 119: 1-6.

Moonlight, P.W., Richardson, J.E., Tebbitt, M.C., Thomas, D.C., Hollands, R., Peng, C.-I & Hughes, M.(2015). Continental-scale diversification patterns in a mega-diverse genus: the biogeography of Neotropical Begonia. Journal of Biogeography DOI: 10.1111/jbi.12496.

Hughes, M. & Takeuchi, W. (2015). A new section (Begonia sect. Oligandrae) and a new species (Begonia pentandra sp. nov.) in Begoniaceae from New Guinea. Phytotaxa, 197(1): 37-44.

Phutthai, T., Hughes, M., Sridith, K. (2014). Begonia kanburiensis (sect. Diploclinium, Begoniaceae), a new species from Thailand. Thai Forest Bulletin 42: 43-47.

Hughes, M. (2013). Memecylaceae. In Kiew et al. (Eds.) Flora of Peninsular Malaysia Series II: Seed Plants. Forestry Research Institute, Malaysia.

Ardi, W.H., Ardhaka, I.M., Hughes, M., Undaharta, N.K.E, Girmansyah, D. & Hidayat, S. (2013). Two new species of Begonia (Begoniaceae) from Bali and Lombok. Gardens' Bulletin Singapore 65(2): 135-142.

Hughes, M. (2013). Memecylon pseudomegarcarpum M.Hughes (Melastomataceae) a new species of tree from Peninsular Malaysia. European Journal of Taxonomy 56: 1-9.

Rubite, R.R., Hughes, M., Alejandro, G.J.D., Peng, C-I. (2013). Recircumscription of Begonia sect. Baryandra (Begoniaceae): evidence from molecular data. Botanical Studies 54: 38.

de Villiers, M.J., Pirie, M.D., Hughes, M. Möller, M., Edwars, T.J., Bellstedt, D.U. (2013). An approach to identify putative hybrids in the 'coalescent stochasticity zone', as exemplified in the African plant genus Streptocarpus (Gesneriaceae). New Phytologist 198(1): 284-300.

Rajbhandary, S., Hughes, M., Shrestha, K.K. (2012) Pollen morphology of Begonia L. (Begoniaceae) in Nepal. Bangladesh Journal of Plant Taxonomy 19(2): 191-200.

Wijedasa, L.S. & Hughes, M. (2012). A new species and new combinations of Memecylon in Thailand and Peninsular Malaysia. Phytotaxa 66: 6-12.

Hughes, M., Girmansyah, D., Handoyo, W.A. & Puglisi, C. (2012). The limestone flora of Sumatra. Gardenwise 39: 18-21.

Hughes, M. (2012). Species old and new: the remarkable Begonia diversity of Sumatra. Begonian, 79(6): 209–214.

Phutthai, T., Hughes, M. & Sridith, K. (2012). A new species of Begonia (Begoniaceae) from Peninsular Thailand. Edinburgh Journal of Botany 69: 287-292.

Thomas, D.C., Hughes, M., Phutthai, T., Ardi, W.H., Rajbhandary, S., Rubite, R., Twyford, A.D., & Richardson, J.E. (2011). West to east dispersal and subsequent rapid diversification of the mega-diverse genus Begonia (Begoniaceae) in the Malesian archipelago. Journal of Biogeography 39: 98-113.

de Wilde, J.J.F.E., Hughes, M., Rodda M. & Thomas D.C. (2011). Pliocene intercontinental dispersal from Africa to Southeast Asia highlighted by the new species Begonia afromigrata(Begoniaceae). Taxon 60: 1685-1692.

Rajbhandary, S., Hughes, M., Phutthai, T., Thomas, D.C. & Shrestha, K.K. (2011). Asian Begonia: out of Africa via the Himalayas? Gardens' Bulletin Singapore 63: 277-286.

Hughes, M. & Girmansyah, D. (2011). Searching for Sumatran Begonia described by William Jack: following in the footsteps of a 19th century Scottish botanist. Gardens' Bulletin Singapore 63: 83-96.

Thomas, D.C., Hughes, M., Phutthai, T., Rajbhandary, S., Rubite, R., Ardi, W.H., Richardson, J. (2011). A non-coding plastid DNA phylogeny of Asian Begonia (Begoniaceae): Evidence for morphological homoplasy and sectional polyphyly. Molecular Phylogenetics and Evolution 60: 428-444.

Thomas, D.C., Ardi, W.H. & Hughes, M. (2011). Nine new species of Begonia L. (Begoniaceae) from South and West Sulawesi, Indonesia. Edinburgh Journal of Botany 68: 225-255.

Hughes, M., Rubite, R.R., Kono, Y. & Peng, C-I. (2011). Begonia blancii (sect. Diploclinium), a new species endemic to the Philippine island of Palawan. Botanical Studies 52: 203-209.

Rajbhandary, S., Hughes, M. & Shrestha, K. (2011). Male flower and androecium morphology of Begonia species in Nepal. Journal of Taxonomy and Biodiversity Research 5: 19-26.

Hughes, M. (2011). A new species of fleshy-fruited Begonia from the Masoala Peninsula, Madagascar. Adansonia 33: 81-85.

Hughes, M., Girmansyah, D. (2011). A revision of Begonia sect. Sphenanthera from Sumatra. Gardens' Bulletin Singapore 62: 239-251.

Rajbhandary, S., Hughes, M. & Shrestha, K. (2010). Distribution patterns of Begonia species in the Nepal Himalaya. Botanica Orientalis, 7:73-78.

Hughes, M., Coyle, C. & Rubite, R.R. (2010). A revision of Begonia section Diploclinium on the Philippine island of Palawan, including five new species. Edinburgh Journal of Botany, 67: 123–140.

Ardi, W.H. & Hughes, M. (2010). Begonia droopiae, a new species from West Sumatra. Gardens' Bulletin Singapore 62: 17-22.

Rajbhandary, S., Hughes, M., & Shrestha, K.K. (2010). Three new species of Begonia sect. Platycentrum from Nepal. Gardens Bulletin Singapore 62: 143-152.

Hughes, M., Girmansyah, D., Ardi, W.H. & Nurainas. (2009). Seven new species of Begonia from Sumatra. Gardens' Bulletin Singapore, 61: 29-44.

Thomas, D.C., Ardi, W.H., Hartutiningsih & Hughes, M. (2009). Two new species of Begonia from South Sulawesi, Indonesia. Edinburgh Journal of Botany, 66: 229-238.

Thomas, D.C., Ardi, W.H. & Hughes, M. (2009). Two new species of Begonia from Central Sulawesi. Edinburgh Journal of Botany, 66: 103-114.

Hughes, M. & Coyle, C. (2009). Begonia section Petermannia on Palawan (Philippines) including two new species. Edinburgh Journal of Botany, 66: 205-211.

Thomas, D.C. & Hughes, M. (2008). Begonia varipeltata (Begoniaceae): A new peltate species from Sulawesi, Indonesia. Edinburgh Journal of Botany, 65: 369-374.

Edwards, T., Hughes, M., Möller, M., Bellstedt, D. (2008). New Streptocarpus species (Gesneriaceae) from South Africa. Botanical Journal of the Linnean Society, 158: 743-748.

Hughes, M. & Hollingsworth, P.M. (2008). Population genetic divergence corresponds with species-level diversity patterns in the large genus Begonia. Molecular Ecology, 17: 2643-2651.

Hughes, M. (2008). An annotated checklist of southeast Asian Begonia. Royal Botanic Garden Edinburgh.

Hughes, M. & Tanaka, N. (2007). A new species of Begonia from northern Myanmar. Acta Phytotaxonomica et Geobotanica, 58: 83-86.

Hughes, M., Möller, M., Bellstedt, D.U., Edwards, T.J. (2007). The impact of pollination syndrome and habitat on gene flow - a comparative study of two Streptocarpus (Gesneriaceae) species. American Journal of Botany, 94: 1688-1695.

Hughes, M. & Pullan, M. (2007).An online database of southeast Asian Begonia. WWW publication:

Hughes, M. (2007). Begonia cladotricha: a new species from Laos. Edinburgh Journal of Botany,64: 102-105.

Hughes, M. (2006). Four new species of Begonia from Sulawesi. Edinburgh Journal of Botany, 63: 191-199.

Hughes, M., Möller, M., Bellstedt, D.U., Edwards, T.J. (2005). Breeding system of a plesiomorphic floral type: an investigation of small flowered Streptocarpus species. Plant Systematics and Evolution, 262: 13-24.

Hughes, M., Möller, M., Bellstedt, D.U., Edwards, T.J & de Villiers, M. (2005). Refugia, dispersal and divergence across a forest archipelago: a study of Streptocarpus in eastern South Africa. Molecular Ecology, 14: 4415-4426.

Forrest, L.L, Hughes, M. & Hollingsworth P.M. (2005). A phylogeny of Begonia using nuclear ribosomal sequence data and non-molecular characters. Systematic Botany, 30: 671-682.

MacMaster, G., Möller, M, Hughes, M., Edwards T.J. & Bellstedt D.U. (2005). A new species of Streptocarpus endemic to Madagacascar. Adansonia, 27: 131-136.

Möller, M., Brooks K.J. & Hughes, M. (2004). Plastid inheritance in Streptocarpus (Gesneriaceae) and an inferred hybrid origin for a population of S. aff. primulifolius from Igoda River, South Africa. Edinburgh Journal of Botany, 60: 389-408.

Hughes, M., Möller, M., Bellstedt, D.U., Edwards, T.J. & Woodhead, M. (2004). EST and random genomic nuclear microsatellites for Streptocarpus. Molecular Ecology Notes, 4: 36-38.

Hughes, M. & Hollingsworth, P.M. (2003). Population structure in the endemic Begonia of the Socotra archipelago. Biological Conservation, 113: 277-284.

Hughes, M., Hollingsworth, P.M. & Squirrell, J. (2002). Isolation of polymorphic microsatellite markers for Begonia sutherlandii Hook. f. Molecular Ecology Notes, 2:185-186.

Hughes, M. & Miller, A.G. (2002). A new endemic species of Begonia (Begoniaceae) from the Socotra archipelago. Edinburgh Journal of Botany, 59: 273-281.

Hughes, M., Russell, J. & Hollingsworth, P.M. (2002). Polymorphic microsatellite markers for the Socotran endemic herb Begonia socotrana. Molecular Ecology Notes, 2: 159-160.

Hughes, M. (2001). Collecting Begonia sutherlandii in Kwazulu Natal, South Africa. GlasgowNaturalist, 23: 125.

Hughes, M. (2001). The Begonia of the Socotra archipelago. The Begonian, 68: 199-213.

Catry, P., Mellanby, R., Suleiman, K.A., Salim, K.H., Hughes, M., McKean, M., Anderson, N., Constant, G., Heany, V., Martin, G., Armitage, M., Wilson, M. (2000). Habitat selection by terrestrial birds on Pemba Island (Tanzania), with particular reference to six endemic taxa. Biological Conservation, 95: 259-267.

Cresswell, W., Hughes, M.,Mellanby, R., Bright, S., Catry, P., Chaves, J., Freile, J., Gabela, A., Martineau, H., MacLeod, R., McPhee, F., Anderson, N., Holt, S., Barabas, S., Chapel, C. & Sanchez T. (1999). Densities and habitat preferences of Andean cloud-forest birds in pristine and degraded habitats in north-eastern Ecuador. Bird Conservation International, 9: 124-145.

Cresswell, W., Mellanby, R., Bright, S., Catry, P., Chaves, J., Freile, J., Gabela, A., Hughes, M.,Martineau, H., MacLeod, R., McPhee, F., Anderson, N., Holt S., Barabas, S., Chapel, C. & Sanchez, T. (1999). The birds of the Guandera Biological Reserve, Carchi, Northern Ecuador. Cotinga, 12: 55-63.

Hughes, M.,Donelly, C., Crozier, A., & Wheeler, C.T. (1999). Effects of the exposure of roots of Alnus glutinosa (L.) Gaertn. to light and flavonoids on nodulation. Canadian Journal of Botany, 77: 1311-1315.

Crozier, A., Hughes, M. & McDonald, M. (1998). Survey of the free and conjugated myricetin and quercetin content of red wines of different geographical origins. Journal of Agricultural and Food Chemistry, 46: 368-375.