Wu X*, Ruhsam M*, Wen Y*, Thomas PI*, Worth JR, Lin X, Wang M, Li X, Chen L, Lamxay V, Le Canh N, Coffman G (2019). The last primary forests of the Tertiary relict Glyptostrobus pensilis contain the highest genetic diversity. Forestry: An International Journal of Forest Research. online early


*These authors contributed equally


One Thousand Plant Transcriptomes Initiative (2019). One thousand plant transcriptomes and the phylogenomics of green plants. Nature, 574(7780), p.679.


Xu J*, Song X*, Ruhsam M, Liu T, Li J, Neaves L E, Miao J, Xie S, Meng Q, Mao K (2019). Distinctiveness, speciation and demographic history of the rare endemic conifer Juniperus erectopatens in the eastern Qinghai-Tibet Plateau. Conservation Genetics, 20(6), 1289-1301.


*These authors contributed equally


Li XY, Lin XY, Ruhsam M, Chen L, Wu XT, Wang MQ, Thomas PI, Wen YF, 2019. Development of microsatellite markers for the critically endangered conifer Glyptostrobus pensilis (Cupressaceae) using transcriptome data. Silvae Genetica, 68(1), 41-44.


Zhou T, Ruhsam M, Wang J, Li W, Zhang X, Xu Y, Xu F, Zhu H, Wang X, 2019. The complete chloroplast genome of Euphrasia regelii, pseudogenization of ndh genes and the phylogenetic relationships within Orobanchaceae. Frontiers in Genetics, 10, 444.


Yan YD, Li XY, Worth JR, Lin XY, Ruhsam M, Chen L, Wu XT, Wang MQ, Thomas PI, Wen, YF (2019). Development of chloroplast microsatellite markers for Glyptostrobus pensilis (Cupressaceae). Applications in Plant Sciences, 7(7), e11277.


Kohn DD, Ruhsam M, Hulme PE, Barrett SCH, Hollingsworth PM (2019). Paternity analysis reveals constraints on hybridization potential between native and introduced bluebells (Hyacinthoides). Conservation Genetics 20(3), 571-584.


Mao K*, Ruhsam M*, Ma Y, Graham SW, Liu J, Thomas P, Milne RI, Hollingsworth PM (2019). A transcriptome-based resolution for a key taxonomic controversy in Cupressaceae. Annals of Botany, 123(1), 153–167.


*These authors contributed equally


Ruhsam M, Jessop W, Cornille A, Renny J, Worrell R. (2018). Crop-to-wild introgression in the European wild apple Malus sylvestris in Northern Britain. Forestry: An International Journal of Forest Research. 92(1), 85-96.


Wang X*, Feng L*, Zhou T*, Ruhsam M, Huang L, Hou X, Song J (2018). Genetic and chemical differentiation characterizes top-geoherb and non-top-geoherb areas in the TCM herb rhubarb. Scientific reports, 8(1), 9424.

*These authors contributed equally

Wang X, Gussarova G, Ruhsam M, de Vere N, Metherell C, Hollingsworth PM, Twyford AD (2018). DNA barcoding a taxonomically complex hemiparasitic genus reveals deep divergence between ploidy levels but lack of species-level resolution. AoB Plants 10(3): ply026

Ruhsam M, PM Hollingsworth (2018). Authentication of Eleutherococcus and Rhodiola herbal supplement products in the United Kingdom. Journal of Pharmaceutical and Biomedical Analysis 149: 403-409.


Mill RR, Ruhsam M, Thomas P, MF Gardner, PM Hollingsworth (2017) Araucaria goroensis (Araucariaceae), a new Monkey Puzzle from New Caledonia, and nomenclatural notes on Araucaria muelleri. Edinburgh Journal of Botany 74(2): 123-139


Ruhsam M, Clark A, Finger A, Wulff AS, Mill RR, Thomas P, Gardner M, Gaudeul M, Ennos RA, PM Hollingsworth (2016). Hidden in plain view: cryptic diversity in the emblematic Araucaria of New Caledonia. American Journal of Botany 103(5): 888-898


Rothfels CJ, Li F-W, Sigel EM, Huiet L, Larsson A, Burge DO, Ruhsam M, Deyholos M, Soltis DE, Stewart CN, Shaw SW, Pokorny L, Chen T, dePamphilis C, DeGironimo L, Chen L, Wei X, Sun X, Korall P, Stevenson DW, Graham SW, Wong GK-S, KM Pryer (2015). The evolutionary history of ferns inferred from 25 low-copy nuclear genes. American Journal of Botany 102: 1089-1107

Ruhsam M, Rai HS, Mathews S, Ross TG, Graham SW, Raubeson LA, Mei W, Thomas P, Gardner MF, Ennos RA, PM Hollingsworth (2015) Does complete plastid genome sequencing improve species discrimination and phylogenetic resolution in Araucaria? Molecular Ecology Resources 15: 1067-1078


Kranitz ML, Biffin E, Clark A, Hollingsworth ML, Ruhsam M, Gardner MF, Thomas P, Mill RR, Ennos RA, Gaudeul M, Lowe AJ, Hollingsworth PM (2014). Evolutionary Diversification of New Caledonian Araucaria. PloS ONE 9: e110308

Li FW, Villarreal JC, Kelly S, Rothfels CJ, Melkonian M, Frangedakis E, Ruhsam M, Sigel EM, Der JP, Pittermann J, Burge DO, Pokorny L, Larsson A, Chen T, Weststrand S, Thomas P, Carpenter E, Zhang Y, Tian Z, Chen L, Yan Z, Zhu Y, Sun X, Wang J, Stevenson DW, Crandall-Stotler BJ, Shaw AJ, Deyholos MK, Soltis DE, Graham SW, Windham MD, Langdale JA, Wong GKS, Mathews S, KM Pryer (2014) Horizontal transfer of an adaptive chimeric photoreceptor from bryophytes to ferns PNAS 111: 6672-6677

Ruhsam M, Jacobs T, Watson K, Hollingsworth PM (2014). Is hybridisation a threat to Rumex aquaticus in Britain? Plant Ecology & Diversity 8(4): 465-474


Rothfels CJ, Larsson A, Li F-W, Sigel EM, Huiet L, Burge DO, Ruhsam M, Graham SW, Stevenson DW, Wong GK-S, Korall P, Pryer KM. (2013) Transcriptome-Mining for Single-Copy Nuclear Markers in Ferns. Plos One 8: e76957

Ruhsam M, Hollingsworth PM, Ennos RA. (2013) Patterns of mating, generation of diversity, and fitness of offspring in a Geum hybrid swarm. Evolution 67: 2728-2740


Ruhsam M, Hollingsworth PM, Ennos RA (2011) Early evolution in a hybrid swarm between outrossing and selfing lineages in Geum. Heredity 107: 246-255


Ruhsam M, Hollingsworth PM, Squirrell J, Ennos RA (2010) Significant differences in outcrossing rate, self-incompatibility and inbreeding depression between two widely hybridising species of Geum. Biological Journal of the Linnean Society. 101: 977-990


Davis AP, Govaerts R, Bridson D, Ruhsam M, Moat J, Brummitt N (2009) A global assessment of distribution, diversity, endemism, and taxonomic effort in the Rubiaceae. Annals of the Missouri Botanical Garden 96: 68-78.


Ruhsam M, Govaerts R, Davis AP (2008) Nomenclatural changes in preparation for a World Rubiaceae Checklist. Botanical Journal of the Linnean Society 157: 115-124.


Ruhsam M, Davis AP (2007) A taxonomic revision of the genus Flagenium Baill. (Rubiaceae-Octotropideae). Botanical Journal of the Linnean Society 155: 557-570.


Davis AP, Ruhsam M (2005). Five new combinations and one new name in Rubiaceae from south-east Asia. Blumea 50: 575-578.


Govaerts R, Ruhsam M, Andersson L, Robbrecht E, Bridson D, Davis A, Schanzer I, Sonké B World Checklist of Rubiaceae. The Board of Trustees of the Royal Botanic Gardens, Kew. Published on the Internet;