Email: David Middleton 
Phone: +44 (0)131 248 2926
Fax: +44 (0)131 248 2901
Research Interests
I am currently working on a revision of the genus Aeschynanthus in the Gesneriaceae and am coordinating the account of the Gesneriaceae for the Flora of Thailand.
My other research interests include Southeast Asian Apocynaceae, subfamilies Rauvolfioideae and Apocynoideae, and general Southeast Asian floristics. I am on the Editorial Boards of the Thai Forest Bulletin and the Flora of Thailand and on the Board of the Flora Malesiana Foundation.
Edinburgh Journal of Botany
I am the editor of the Edinburgh Journal of Botany. The Edinburgh Journal of Botany (EJB) is an international journal of plant systematics covering related aspects of biodiversity, conservation science and phytogeography. Published by Cambridge University Press in partnership with the Royal Botanic Garden Edinburgh, the journal provides global coverage of all groups of plants and fungi.
Publications
1. MIDDLETON, D.J. & WILCOCK, C.C. (1990). A critical examination of the status of Pernettya Gaud. as a genus distinct from Gaultheria L. Edinburgh Journal of Botany 47: 291-301
2. MIDDLETON, D.J. & WILCOCK, C.C. (1990). Chromosome counts in the genus Gaultheria L. and related genera. Edinburgh Journal of Botany 47: 303-313
3. MIDDLETON, D.J. (1991). Pernettya or Gaultheria? The Plantsman 20:167-177
4. MIDDLETON, D.J. (1991). Ecology, reproductive biology and hybridisation in the genus Gaultheria L. (Ericaceae). Edinburgh Journal of Botany 48: 81-89
5. MIDDLETON, D.J. (1991). Infrageneric classification of the genus Gaultheria L. (Ericaceae). Botanical Journal of the Linnean Society 106: 229-258
6. MIDDLETON, D.J. (1991). Taxonomic studies in the Gaultheria group of genera of the tribe Andromedeae (Ericaceae). Edinburgh Journal of Botany 48: 283-306
7. MIDDLETON, D.J. (1991). Gaultheria x wisleyensis: a not-so-new hybrid! The Plantsman 13: 188-189
8. MIDDLETON, D.J. (1992). A chemotaxonomic survey of flavonoids and simple phenols in the leaves of Gaultheria L. and related genera. Botanical Journal of the Linnean Society 110: 313-324
9. MIDDLETON, D.J. (1993). A taxonomic revision of Willughbeia Roxb. (Apocynaceae). Blumea 38: 1-24
10. MIDDLETON, D.J. (1993). A systematic survey of leaf and stem anatomical characters in the genus Gaultheria and related genera (Ericaceae). Botanical Journal of the Linnean Society 113: 199-215
11. MIDDLETON, D.J. (1993). A new combination in Chonemorpha (Apocynaceae). Novon 3: 455
12. MIDDLETON, D.J. (1994). A new combination in Epigynum (Apocynaceae). Kew Bulletin 49: 304
13. MIDDLETON, D.J. (1994). New combinations in Urceola (Apocynaceae). Novon 4: 151
14. MIDDLETON, D.J. (1994). Three new combinations in Chinese Apocynaceae. Novon 4: 152
15. MIDDLETON, D.J. & GILBERT, M. (1994). (1104) Proposal to reject Alyxia glaucescens Wall. Taxon 43: 477-478
16. MIDDLETON, D.J. (1994). A taxonomic revision of Ichnocarpus (Apocynaceae). Blumea 39: 73-94
17. MIDDLETON, D.J. (1995). A revision of the genus Urceola (Apocynaceae) in Thailand. Kew Bulletin 49: 757-767
18. MIDDLETON, D.J. (1995). Two new species of Alyxia (Apocynaceae: Plumerioideae) from Thailand. Blumea 40: 109-111
19. MIDDLETON, D.J. (1995). Baharuia, a new genus of Apocynaceae from Malesia. Blumea 40: 443-447
20. MIDDLETON, D.J. (1995). A revision of Papuechites (Apocynaceae). Blumea 40: 439-442
21. SIMPSON, D.A., PARNELL, J.A.N., CHANTARANOTHAI, P. & MIDDLETON, D.J. (1995). The Royal Botanic Gardens Kew, Khon Kaen University and Trinity College Dublin expeditions to Thailand 1990 and 1993. Thai Forest Bulletin (Bot.) 23: 50-61
22. LI, P.T., LEEUWENBERG, A.J.M. & MIDDLETON, D.J. (1996). Apocynaceae in Flora of China 16: 143-188
23. MIDDLETON, D.J. (1996). Ecua, a new genus of Apocynaceae from Malesia. Blumea 41: 33-35
24. MIDDLETON, D.J. (1996). A revision of Anodendron Roxb. (Apocynaceae). Blumea 41: 37-68
25. MIDDLETON, D.J. (1996). A revision of Aganonerion Pierre ex Spire, Parameria Benth. and Urceola Roxb. (Apocynaceae). Blumea 41: 69-122
26. MIDDLETON, D.J. (1996). A new species of Willughbeia (Apocynaceae: Plumerioideae) from Borneo. Blumea 41: 123-124
27. MIDDLETON, D.J. (1996). A revision of Aganosma (Bl.) G.Don (Apocynaceae). Kew Bulletin 51: 455-482
28. MIDDLETON, D.J. (1997). (1280) A proposal to reject Apocynum vincifolium Burm.f. so as to retain Parsonsia alboflavescens (Dennst.) Mabberley. Taxon 46: 121-122
29. MIDDLETON, D.J. (1997). A revision of Parsonsia R.Br. in Malesia. Blumea 42: 191-248.
30. MIDDLETON, D.J. (1997). A revision of Carruthersia Seem. Blumea 42: 489-498
31. MIDDLETON, D.J. (1997). Gaultheria. European Garden Flora 5: 492-499
32. MIDDLETON, D.J. (1997). Profile - Max van Balgooy. Plant Talk 11: 7
33. MIDDLETON, D.J. (1997). Accounts of several families in G. Argent, A. Saridan, E. Campbell, P. Wilkie. (Eds). Manual of the larger and more important non dipterocarp trees of Central Kalimantan.
34. ZIJLSTRA, G., LEEUWENBERG, A.J.M & MIDDLETON, D.J. (1998). (1332) Proposal to conserve the name Trachelospermum (Apocynaceae) with a conserved type. Taxon 47: 163-164
35. MIDDLETON, D.J. & LEEUWENBERG, A.J.M. (1998). (1355) Proposal to conserve Tabernaemontana heyneana Wall. (Apocynaceae). Taxon 47: 481-482
36. MIDDLETON, D.J. (1999). Apocynaceae. Flora of Thailand 7(1): 1-152
37. HENDRIAN & MIDDLETON, D.J. (1999). A revision of Rauvolfia L. in Malesia. Blumea 44: 449-470
38. MIDDLETON, D.J. (2000). Revision of Alyxia (Apocynaceae). Part 1: Asia and Malesia. Blumea 45: 1-146
39. MIDDLETON, D.J. (2001). A new species of Pottsia (Apocynaceae: Apocynoideae) from Thailand and Lao PDR. Harvard Papers in Botany 6: 285-287
40. BEAMAN, J.H. & MIDDLETON, D.J. (2001). Apocynaceae, pp. 102-114 in Beaman, J. H., Anderson, C., & Beaman, R. S. Plants of Mount Kinabalu, 4. Dicotyledon Families Acanthaceae to Lythraceae.
41. MIDDLETON, D.J. & SANTISUK, T. (2001). A new species of Wrightia (Apocynaceae: Apocynoideae) from Thailand. Thai Forest Bulletin 29: 1-10
42. MIDDLETON, D.J. (2001). The Apocynaceae of the Crocker Range National Park. In: Ghazaly, I. L. & A. Lamri (eds.), Crocker Range National Park Sabah, vol. 1. Natural Ecosystem and species components, pp. 65-87. Asean Academic Press, London.
43. SANGAT-ROEMANTYO, H.M. & MIDDLETON, D.J. (2001). Alyxia R.Br. In: Van Valkenburg, J.L.C.H. & Bunyapraphatsara, N. (Eds.), Plant Resources of South-East Asia, vol. 12(2), Medicinal and poisonous plants 2: 69-72
44. MIDDLETON, D.J. (2002). Revision of Alyxia (Apocynaceae). Part 2: Australia and Pacific Islands. Blumea 47: 1-93
45. MIDDLETON, D.J. (2002). Trachelospermum axillare Hook.f. (Apocynaceae), a new record for Thailand. Thai Forest Bulletin 30: 28-30
46. PARNELL, A.N., SIMPSON, D.A., MOAT, J., KIRKUP, D.W., CHANTARANOTHAI, P., BOYCE, P.C., BYGRAVE, P., DRANSFIELD, S., JEBB, M. H. P., MACKLIN, J., MEADE, C., MIDDLETON, D.J., MUASYA, A. M., PRAJAKSOOD, A., PENDRY, C.A., POOMA, R., SUDDEE, S., & WILKIN, P. (2003). Plant collecting spread and densities: their potential impact on biogeographical studies in Thailand. Journal of Biogeography 30: 193-209
47. MIDDLETON, D.J. (2003). Progress on the Flora of Thailand. Telopea 10: 33-42
48. MIDDLETON, D.J. (2003). A new species of Carruthersia (Apocynaceae: Apocynoideae) from the Philippines. Harvard Papers in Botany 8: 1-3
49. MIDDLETON, D.J. (2003). A new species and a new combination in Bornean Kopsia (Apocynaceae: Rauvolfioideae). Gardens Bulletin Singapore 55: 65-68
50. LINDSAY, S. & MIDDLETON, D.J. (2003). Proposal to conserve the name Davallia repens (L. f.) Kuhn (Davalliaceae) against Davallia repens (Bory) Desv. (Lindsaeaceae). Taxon 52: 630-631
51. MIDDLETON, D.J. (2003). A revision of Dyera (Apocynaceae, Rauvolfioideae). Gardens Bulletin Singapore 55: 209-217
52. LINDSAY, S., SUDDEE, S., MIDDLETON, D.J. & POOMA, R. (2004). Matoniaceae (Pteridophyta) - a new family record for Thailand. Thai Forest Bulletin 31: 47-52
53. SUDDEE, S. & MIDDLETON, D.J. (2004). Rhodoleia (Hamamelidaceae), a new generic record for Thailand. Thai Forest Bulletin 31: 132-135
54. LINDSAY, S. & MIDDLETON, D.J. (2004). Adiantum phanomensis (Adiantaceae), a new fern species from Peninsular Thailand. Harvard Papers in Botany 8: 137-140
55. MIDDLETON, D.J. (2004). A revision of Kopsia Blume (Apocynaceae: Rauvolfioideae). Harvard Papers in Botany 9: 91-144
56. MIDDLETON, D.J. (2004). Apocynaceae. Tree Flora of Sabah and Sarawak vol. 5: 1-61
57. RUNDEL, P.W., MIDDLETON, D.J., PATTERSON, M. & MENG, M. (2004) [‘2003']. Structure and ecological function in a tropical montane Sphagnum bog of the Elephant Mountains, Bokor National Park, Cambodia. Natural History Bulletin of the Siam Society 51(2): 185-196
58. MIDDLETON, D.J. (2004). Proposal to reject the name Echites trichotoma (Apocynaceae: Apocynoideae). Taxon 53: 1071-1072
59. BOONKERD, T., LINDSAY, S., MIDDLETON, D.J. & SUDDEE, S. (2004). Additions to the Pteridophyte flora of Thailand. Thai Forest Bulletin 32: 6-11
60. MIDDLETON, D.J. (2005). A new species of Tabernaemontana (Apocynaceae: Rauvolfioideae) from the Philippines. Harvard Papers in Botany 9: 387-389
61. MIDDLETON, D.J. (2005). A new species of Micrechites (Apocynaceae: Apocynoideae) from Borneo. Harvard Papers in Botany 9: 383-385
62. MIDDLETON, D.J. (2005). A new species of Alstonia (Apocynoideae: Rauvolfioideae) from Vietnam. Harvard Papers in Botany 10: 63-65
63. MIDDLETON, D.J. (2005). A revision of Epigynum (Apocynaceae: Apocynoideae). Harvard Papers in Botany 10: 67-81
64. MIDDLETON, D.J. (2005). A revision of Wrightia (Apocynaceae: Apocynoideae) in Malesia. Harvard Papers in Botany 10: 161-182
65. MIDDLETON, D.J. (2005). A new species of Kopsia (Apocynoideae: Rauvolfioideae) from Vietnam. Adansonia 27: 287-289
66. MIDDLETON, D.J. & REGALADO, J. (2005). Nomenclatural notes on species of Apocynaceae described from Cambodia, Laos and Vietnam. Adansonia 27: 291-308
67. MIDDLETON, D.J., LINDSAY, S. & SUDDEE, S. (2006 [‘2005']). A new species of Kamettia (Apocynaceae: Rauvolfioideae), a genus new to Thailand. Thai Forest Bulletin 33: 75-80
68. NGERNSAENGSARUAY, C., MIDDLETON, D.J. & CHAYAMARIT, K. (2006). Five new records of Litsea (Lauraceae) for Thailand. Thai Forest Bulletin 33: 81-93
69. MIDDLETON, D.J. (2006). Nomenclatural notes on Asian Apocynaceae, subfamilies Rauvolfioideae and Apocynoideae. Taxon 55: 502-506
70. MIDDLETON, D.J. (2006). Three new species of Chilocarpus (Apocynaceae: Rauvolfioideae) from Malesia. Edinburgh Journal of Botany 63: 201-207.
71. MIDDLETON, D.J. (2007). A new species and a new combination in Aeschynanthus (Gesneriaceae) from Laos. Edinburgh Journal of Botany 64: 45-50
72. MIDDLETON, D.J. (2007). Apocynaceae, subfamilies Rauvolfioideae and Apocynoideae. Flora Malesiana 18: 1-471
73. LIVSHULTZ, T., MIDDLETON, D.J., ENDRESS, M.E. & WILLIAMS, J. (2007). Phylogeny of Apocynoideae and the APSA clade. Annals of the Missouri Botanical Garden 94: 323-361
74. MIDDLETON, D.J. (2007). Amphineurion (Apocynaceae) and Amphineuron (Pteridophyta). Taxon 56: 593
75. MIDDLETON, D.J. (2007). Angiosperms: Apocynaceae of Papua. In: Marshall, A.J. & Beehler, B.M., The Ecology of Papua, vol. 1: 355-358
76. MIDDLETON, D.J. (2007). A revision of Aeschynanthus (Gesneriaceae) in Thailand. Edinburgh Journal of Botany 64: 363-429
77. CHAYAMARIT, K., CHANTARANOTHAI, P., MIDDLETON, D.J., PARNELL, J.A.N., SIMPSON, D.A. & WILKIN, P. (2007). What constitutes a new taxon record for Thailand? Thai Forest Bulletin 35: 1-3
78. MIDDLETON, D.J. (2007). A new species of Wrightia (Apocynaceae: Apocynoideae) from Thailand. Thai Forest Bulletin 35: 80-85
79. MIDDLETON, D.J. (2007). Chilocarpus rostratus (Apocynaceae: Rauvolfioideae), a new record for Thailand. Thai Forest Bulletin 35: 86-88
80. MIDDLETON, D.J. & SCOTT, S.M. (2008). A new species of Agalmyla (Gesneriaceae) from Sulawesi. Edinburgh Journal of Botany 65: 49-52
81. XU, Z.R., BURTT, B. L., SKOG, L.E. & MIDDLETON, D.J. (2008). A Revision Of Paraboea (Gesneriaceae). Edinburgh Journal of Botany 65: 161-347